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Grey Seal - 2 bulls fighting in surf during breeding season. Donna Nook seal sanctuary, Lincolnshire, UKUSH-2552 Grey Seal - 2 bulls fighting in surf during breeding season Donna Nook seal sanctuary, Lincolnshire, UK Halichoerus grypus Duncan Usher Please note that prints are for personal display
European Hare - x2 boxingUSH-202 European HARES - boxing Lepus europaeus Duncan Usher Please note that prints are for personal display purposes only and may not be reproduced in any way
European Wolf - young wolf confronting rank weakest animal, pack dispute Bavaria, GermanyUSH-2006 European Wolf - young wolf confronting rank weakest animal, pack dispute. Note tail position is high of dominant wolf and low on the submissive wolf
European Wolf - Young animal attacking the rank weakest animal in snow, winter Bavaria, GermanyUSH-2002 European Wolf - Young animal attacking the rank weakest animal in snow, winter Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only
European Wolf - pack showing aggresssion towards rank weakest animal Bavaria, GermanyUSH-1975 European Wolf - pack showing aggresssion towards rank weakest animal Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only
Burchell's / Plains / Common Zebra - stallions, Dominance behavior. Serengeti National Park, Tanzania. May. 3MB778TOM-864 Burchell's / Plains / Common Zebra - stallions, dominance behavior. Serengeti National Park, Tanzania. May. Equus burchelli Tom & Pat Leeson Please note that prints are for personal
Masai Giraffe - Young males "necking" (dominance behavior). East Africa 3mb496TOM-861 Masai Giraffe - Young males "necking" (dominance behavior). Kenya, Africa Giraffa camelopardalis tippelski Tom & Pat Leeson Please note that prints are for personal display purposes
Bighorn Sheep - rams butt heads Northern Rockies. Fall. MS363TOM-1589 Bighorn Sheep - rams butt heads Yellowstone National Park, Northern Rockies. USA, Fall. Ovis canadensis Tom & Pat Leeson Please note that prints are for personal display purposes only
Northern or Steller Sea Lions - bull guarding territory and harem. Pacific Northwest. ML479TOM-1547 Northern / Steller Sealions - bull guarding territory and harem Pacific Northwest, North America Eumetopias jubatus Tom & Pat Leeson Please note that prints are for personal display purposes
sage grouse - two male slap wings (fight) over territory on a lek during spring mating rituals. Western U. S. A B6015TOM-1485 Sage grouse - two male slap wings (fight) over territory on a lek during spring mating rituals. Western U.S.A Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for
Sage Grouse - two male fight over territory on a lek during spring mating rituals. Western U. S. A. BG193TOM-1482 Sage Grouse - two male fight over territory on a lek during spring mating rituals Western U.S.A. Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for personal display
Sage Grouse - two male confront one another over territory on a lek during spring mating season. Western U. S. A. BG313TOM-1481 Sage Grouse - two male confront one another over territory on a lek during spring mating season. Western U.S.A. Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for
Coastal Grizzly Bears wrestling - dominance behavior among big males at salmon fishing areas, McNeil River, AlaskaTOM-1479 Coastal Grizzly Bears wrestling - dominance behavior among big males at salmon fishing areas McNeil River, Alaska
Mountain Goat - two males in a bluff fight, common dominance behavior between goats. Pacific Northwest. MG57TOM-1384 Rocky Mountain Goat - two males in a bluff fight, common dominance behavior between goats Washington, Pacific Northwest
Pronghorn - buck and doe. Buck is chasing doe who has bolted from his harem. Yellowstone National Park, Montana, USATOM-1360 Pronghorn - buck and doe. Buck is chasing doe who has bolted from his harem. Yellowstone National Park, Montana, USA
Bison - bulls sparring Western U. S. A. MB468TOM-1337 Bison - bulls sparring Theodore Roosevelt National Park, North Dakota, U.S.A. Bison bison Tom & Pat Leeson Please note that prints are for personal display purposes only
Bull Bison - fighting, sparring, summer. Theodore Roosevelt National Park, North Dakota. MB249TOM-1335 Bull Bison - fighting, sparring, summer Theodore Roosevelt National Park, North Dakota, USA Bison bison Tom & Pat Leeson Please note that prints are for personal display purposes only
Grey Wolves In winter involved in dominance behavior--usually no one gets hurt in these short but intense displaysTOM-1299 Grey Wolves In winter involved in dominance behavior - usually no one gets hurt in these short but intense displays Minnesota
Gray Wolves (Canis lupus) displaying dominance over wolf on ground. In snow, MontanaTOM-1294 Grey Wolves - displaying dominance over wolf on ground. In snow Montana Canis lupus Tom & Pat Leeson Please note that prints are for personal display purposes only
Gray Wolves (Canis lupus) in snow, involved in dominance behavior - usually no one gets hurt in these short but intenseTOM-1286 Grey Wolves - in snow, involved in dominance behavior - usually no one gets hurt in these short but intense displays Minnesota
Grey Wolf / Timber Wolf - Two, dominance behavior in snow. montanaTOM-1284 Grey Wolf / Timber Wolf - Two, dominance behavior in snow Montana, USA Canis lupus Tom & Pat Leeson Please note that prints are for personal display purposes only
Grey Wolf / Timber Wolf - dominance behavior. Minnesota, USATOM-1276 Grey Wolf / Timber Wolf - dominance behavior. Minnesota, USA Canis lupus Captive in a reserve Tom & Pat Leeson Please note that prints are for personal display purposes only
Grey Wolf / Timber Wolf - dominance behavior in snowTOM-1271 Grey Wolf / Timber Wolf - dominance behavior in snow Canis lupus Tom & Pat Leeson Please note that prints are for personal display purposes only and may not be reproduced in any way
Bighorn Sheep - rams butt heads. Jasper National Park, Rocky Mountains, Alberta, Canada. MS334TOM-1264 Bighorn Sheep - rams butt heads Jasper National Park, Rocky Mountains, Alberta, Canada Tom & Pat Leeson Please note that prints are for personal display purposes only
Coastal Grizzly Bears wrestling. Nr McNeil River, Alaska. Summer MA1305TOM-1221 Coastal Grizzly Bears - males wrestling. Nr McNeil River, Alaska. Summer Ursus arctos horribilis Tom & Pat Leeson Please note that prints are for personal display purposes only
California Bighorn Sheep - rams dominance behaviorTOM-113 California Bighorn SHEEP- Rams, dominance behavior October Orvis canadensis californiana Ovis canadensis californiana Tom & Pat Leeson Please note that prints are for personal display
Black Bears (Ursus americanus) fighting--mostly snapping jaws and growling--over fishing spot. S. E. Alaska. SummerTOM-1105 Black Bears - fighting, mostly snapping jaws and growling over fishing spot S.E. Alaska. Summer Ursus americanus Tom & Pat Leeson Please note that prints are for personal display purposes
Rocky Mountain Bighorn Sheep - rams. Dominance behavior. Yellowstone National Park, Rockies, Western USA. Autumn MS253TOM-1047 Rocky Mountain Bighorn / Big-Horn SHEEP - rams. Dominance behavior. Yellowstone National Park, Rockies, Western USA
WolfSM-2189 Timber Wolf - intimidation (priority behaviour), Germany canis lupus lycaon Stefan Meyers Please note that prints are for personal display purposes only and may not be reproduced in any way
WolfSM-2188 Wolf - intimidation (priority behaviour) Germany canis lupus lycaon Stefan Meyers Please note that prints are for personal display purposes only and may not be reproduced in any way
European Brown Bears - Two together in snow being aggressiveSM-1910 European Brown Bears - Two together in snow being aggressive Ursus arctos Stefan Meyers Please note that prints are for personal display purposes only and may not be reproduced in any way
Racoon - aggresive/submissive behaviourSM-1519 Raccoon - aggresive / submissive behaviour Stefan Meyers Please note that prints are for personal display purposes only and may not be reproduced in any way
Lion adult male reprimanding a young one Etosha National Park, Namibia, AfricaSAS-63 Lion - adult male reprimanding a young one Etosha National Park, Namibia, Africa Panthera leo Steffen & Alexandra Sailer Please note that prints are for personal display purposes only
Hooker's sealion - on beach guarding territory. Serat Bay Catlins - South Island - New ZealandMAB-455 Hooker's sealion - on beach guarding territory Serat Bay Catlins - South Island - New Zealand. Phocarctos hookeri This is one of the rarest
Black-backed Jackals Two fighting Maasai Mara, AfricaLA-827 Black-backed Jackals - Two fighting with Hyaena in background feeding at kill Maasai Mara, Africa Canis mesomelas Jean Michel Labat Please note that prints are for personal display purposes
European Brown Bear - two playingLA-1677 European Brown Bear - two playing Ursus arctos Jean Michel Labat Please note that prints are for personal display purposes only and may not be reproduced in any way
Young Royal Bengal Tigers play-fighting, Ranthambhor National Park, IndiaJR-1016 Young Royal Bengal Tigers play-fighting Ranthambhore National Park, India Panthera tigris Alternative spellings: Ranthambhor / Ranthambore / Ranthambor Sequence
Bengal / Indian Tigers - two young play fightingJR-1014 Bengal / Indian Tigers - two young play fighting Ranthambhore National Park, India. Panthera tigris Alternative spellings: Ranthambhor / Ranthambore / Ranthambor Sequence
Ibex - fightingJPF-4201 European / Alpine Ibex - fighting on cliff edge Gran Paradiso National Park, Italy Capra hircus ibex Jean Paul Ferrero Please note that prints are for personal display purposes only
Southern Elephant Seal - Dominant male; female. Lesser males further back on the beachFG-dz-877 Southern Elephant Seal - Dominant male; female. Lesser males further back on the beach. Valdes Peninsula, Patagonia
Northern Elephant Seal - males fighting Central California, USAFG-10311 Northern Elephant Seal - males fighting Central California, USA. Mirounga angustirostris Francois Gohier Please note that prints are for personal display purposes only
Topi Maasai Mara, Kenya, AfricaED-292 Korrigum - two males sparring. Maasai Mara, Kenya, Africa Damaliscus lunatus korrigum Korrigum is a subspecies of the Topi, D
African Wild Dogs - Young Wild Dogs mock fighting Moremi, Botswana, AfricaCRH-1002 African Wild Dogs - Young Wild Dogs mock fighting Moremi, Botswana, Africa Lycaon pictus Chris Harvey Please note that prints are for personal display purposes only
Robins – fighting West Wales UK 003185BB-1163 Robins - fighting West Wales UK Erithacus rubecula Brian Bevan Please note that prints are for personal display purposes only and may not be reproduced in any way
Robin – attacking china model West Wales UK 003178BB-1162 Robin - attacking china model West Wales UK Erithacus rubecula Brian Bevan Please note that prints are for personal display purposes only and may not be reproduced in any way
Lesser Kestrel - two males fighting. Caceres - Extramadura - SpainWAT-12222 Lesser Kestrel - two males fighting Caceres - Extramadura - Spain Falco naumanni M. Watson Please note that prints are for personal display purposes only
Greater Flamingo - in water, aggressive posture. Saintes Maries de la Mer - CamargueWAT-11943 Greater Flamingo - in water, aggressive posture Saintes Maries de la Mer, Camargue, France M. Watson Please note that prints are for personal display purposes only
Cuttlefish courting - Two male cuttlefish courting a female. both males struggled with each other to hold the femaleVT-8364 Cuttlefish courting - Two male cuttlefish courting a female. both males struggled with each other to hold the female Thailand, Andaman Sea, Richlieu Rock