Social Behaviour Collection

Wolf / Gray Wolf / Timber Wolf - submissive behaviour Minnesota USA
WAT-14126 Wolf / Gray Wolf / Timber Wolf - submissive behaviour Minnesota USA Canis lupus M. Watson Please note that prints are for personal display purposes only

Blue & Yellow Macaws x2 preening. Taman Burning Bali Bird Park, Bali Island, Indonesia
ANZ-618 Blue and Yellow MACAWS / Blue and Gold macaws - X2 preening Taman Burning Bali Bird Park, Bali Island, Indonesia. Ara ararauna Andrey Zvoznikov Please note that prints are for personal

Japanese Macaque Monkeys - in water - Joshinetsu Kogen NP - Honshu - Japan

JPF-10209 Japanese Macaque Monkeys - in water Joshinetsu Kogen NP - Honshu - Japan Macaca fuscata Jean Paul Ferrero Please note that prints are for personal display purposes only

Two Grey Wolves playing - dominance behavior. Note: there is a third wolf whose ears just show at the bottom of
TOM-1270 Two Grey Wolves playing - dominance behavior Canis lupus Tom & Pat Leeson Please note that prints are for personal display purposes only and may not be reproduced in any way

Sulphur Crested Cockatoos - two birds preening - Victoria Dandenong -Melbourne - Australia
BMD-216 Sulphur Crested Cockatoos - two birds preening Victoria Dandenong -Melbourne - Australia Cacatua sulphurea Brian McDairmant Please note that prints are for personal display purposes only

Gelada Baboon - showing gums & teeth. Simien mountains - Ethiopia - Africa
WAT-10013 Gelada Baboon - male showing gums & teeth Simien mountains - Ethiopia - Africa Theropithecus gelada M. Watson Please note that prints are for personal display purposes only

European Wolf- male and female alpha wolf attacking rival male, in snow Bavaria, Germany
USH-1995 European Wolf - male and female alpha wolf attacking rival male, in snow Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only

African Spoonbill - mutual preening
PS-3702 African Spoonbill - mutual preening Platalea alba Peter Steyn contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

Olive Baboon RS 210 Grooming young, social bonding Papio cynocephalus © Robyn Stewart / ardea. com
RS-210 Olive BABOON - grooming young; social bonding Papio cynocephalus Robyn Stewart contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

European Wolf USH 288 Threatening behaviour Canis lupus © Duncan Usher / ardea. com
USH-288 European Wolf - threatening behaviour captive Canis lupus Duncan Usher contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

Impala - pair involved in social grooming - Kruger National Park - South Africa
VAN-31 IMPALA - Young pair involved in social grooming Krugar National Park, South Africa Aepyceros melampus Ingrid Van den Berg contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

Lions sparring CRH 914 Two young male lions sparring out on floodplains Moremi
CRH-914 Lions sparring - Two young male lions sparring out on floodplains Moremi, Botswana Panthera leo Chris Harvey contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

American White Pelican - feeding together
WW-258 American White Pelican - feeding together Pelecanus erythrorhynchos Wardene Weisser contact details: prints@ardea.com tel: +44 (0) 20 8318 1401

Alaskan / Northern Sea Otter - raft - Sea Otters often gather in a protected area (from current tide wind & waves)
TOM-1920 Alaskan / Northern Sea Otter - raft - Sea Otters often gather in a protected area (from current tide wind & waves)

Olive Baboon - Adult females grooming (with infant) Gombe Stream Reserve, Tanzania
SE-764 Olive Baboon - Adult females grooming (with infant) Gombe Stream Reserve, Tanzania Papio anubis Suzi Eszterhas Please note that prints are for personal display purposes only

Mountain Gorilla - Social grooming. Virunga Volcanoes National Park - Rwanda. Endangered Species
SE-553 Mountain Gorilla - Social grooming. Virunga Volcanoes National Park - Rwanda. Endangered Species Gorilla gorilla berengei Suzi Eszterhas Please note that prints are for personal display

Black-browed Albatross - breeding pair grooming Steeple Jason, Falkland Islands
SE-447 Black-browed Albatross - breeding pair grooming Steeple Jason, Falkland Islands Thalassarche melanophris Suzi Eszterhas Please note that prints are for personal display purposes only

Black-backed Jackal - 16 week old pups grooming each other. Maasai Mara Triangle - Kenya
SE-650 Black-backed Jackal - 16 week old pups grooming each other. Maasai Mara Triangle - Kenya Canis mesomelas Suzi Eszterhas Please note that prints are for personal display purposes only

Mustangs / Wild Horses - standing in meadow carpeted with wildflowers - Western U. S. - Summer _D3C9239
TOM-1908 Mustangs / Wild Horses - standing in meadow carpeted with wildflowers Western U.S. - Summer Equus ferus caballus Tom & Pat Leeson Please note that prints are for personal display purposes

Barbary Macaque / Ape - grooming - Gibraltar - Europe
RD-127 Barbary Macaque / Ape - grooming Gibraltar - Europe Macaca sylvanus Giacomo Radi Please note that prints are for personal display purposes only and may not be reproduced in any way

Japanese Macaque - adult grooming youngster on pool edge - Jigokudani Park - Japan
GET-2095 Japanese Macaque - adult grooming youngster on pool edge Jigokudani Park - Japan Macaca fuscata Geoff Trinder Please note that prints are for personal display purposes only

Barbary Ape JPF 6970 Grooming Macaca sylvana © Jean-Paul Ferrero / ARDEA LONDON
JPF-6970 Barbary Macaque / Barbary Ape / Rock Ape - Grooming Macaca sylvana Calling this species an ape is misleading, for though it lacks a tail

Rhesus Macaque Monkey - cleaning mate, Keoladeo, National Park, India
JR-211 Rhesus Macaque MONKEY- Cleaning mate Keoladeo, National Park, India Macaca mulatta Jagdeep Rajput Please note that prints are for personal display purposes only

Japanese Macaque - pair of adults grooming - Jigokudani Park - Japan
GET-2105 Japanese Macaque - pair of adults grooming Jigokudani Park - Japan Macaca fuscata Geoff Trinder Please note that prints are for personal display purposes only

Japanese Macaque - pair of adults grooming sitting on rock ledge - Jigokudani Park - Japan
GET-2106 Japanese Macaque - pair of adults grooming sitting on rock ledge Jigokudani Park - Japan Macaca fuscata Geoff Trinder Please note that prints are for personal display purposes only

Chimpanzee - mother grooming sub-adult offspring (with three month old infant in lap)
SE-58 Chimpanzee - mother grooming sub-adult offspring (with three month old infant in lap) tropical forest - Western Uganda - Africa Pan troglodytes Suzi Eszterhas Please note that prints are for

Chimpanzee - sub-adult grooming three year old sibling - tropical forest - Western Uganda - Africa
SE-49 Chimpanzee - sub-adult grooming three year old sibling tropical forest - Western Uganda - Africa Pan troglodytes Suzi Eszterhas Please note that prints are for personal display purposes only

Chimpanzee - family group showing social grooming behavior - tropical forest - Western Uganda - Africa
SE-56 Chimpanzee - family group showing social grooming behaviour tropical forest - Western Uganda - Africa Pan troglodytes Suzi Eszterhas Please note that prints are for personal display purposes

Chimpanzee - social grooming - Ngamba Island Chimpanzee Sanctuary
SE-83 Chimpanzee - social grooming Ngamba Island Chimpanzee Sanctuary, Uganda Pan troglodytes Suzi Eszterhas Please note that prints are for personal display purposes only

Chimpanzee - social grooming in morning sun - tropical forest - Western Uganda - Africa
SE-19 Chimpanzee - social grooming in morning sun tropical forest - Western Uganda - Africa Pan troglodytes Suzi Eszterhas Please note that prints are for personal display purposes only

Barbary Macaque - group mutual grooming 
JPF-6964 Barbary Macaque / Barbary Ape / Rock Ape Macaca sylvana - group grooming Macaca sylvanus Calling this species an ape is misleading, for though it lacks a tail

Black-tailed Prairie Dog Polar Park, Canada
ED-389 Black-tailed Prairie Dog Polar Park, Canada Cynomys ludovicianus Eric Dragesco Please note that prints are for personal display purposes only and may not be reproduced in anyway

Barbary Macaques grooming young
DOW-170 Barbary Macaques grooming young. Macaca sylvanus Steve Downer Please note that prints are for personal display purposes only and may not be reproduced in anyway

Barbary Macaques grooming
DOW-172 Barbary Macaques grooming. Macaca sylvanus Steve Downer Please note that prints are for personal display purposes only and may not be reproduced in anyway

Ape: Mountain Gorillas - juveniles playing, Virunga Volcanoes, Rwanda, Africa
AW-6795 Ape: Mountain Gorillas - juveniles playing Virunga Volcanoes, Rwanda, Africa Gorilla gorilla beringei Adrian Warren Please note that prints are for personal display purposes only

Wild Grey Wolf - approximately 6 month old pup on left playing with mother - Autumn - Greater Yellowstone Ecological
TOM-1710 Wild Grey Wolf - approximately 6 month old pup on left playing with mother - Autumn Greater Yellowstone Ecological Area - Yellowstone National Park - Wyoming - USA Canis lupus Tom & Pat

Wild Grey Wolves - greeting in snow - closer wolf is approximately 6 months old pup greeting his mother - Greater
TOM-1693 Wild Grey Wolves - greeting in snow - closer wolf is approximately 6 months old pup greeting his mother Greater Yellowstone Ecological Area - Wyoming - USA Canis lupus Tom & Pat Leeson

Wild Grey Wolf - in snow - approximately 6 month old pup on left showing submission /playfulness to adults
TOM-1691 Wild Grey Wolf - in snow - approximately 6 month old pup on left showing submission /playfulness to adults (creamier wolf is his mother)

Pygmy Chimpanzee - grooming
KFO-989 PYGMY / BONOBO CHIMPANZEES - mutual grooming x five Pan paniscus Kenneth W Fink Please note that prints are for personal display purposes only and may not be reproduced in any way

Chimpanzee KEL 1078 Grooming. Forests, broken savannas of Senegal, Guinea & Cameroon
KEL-1078 Chimpanzee - adult grooming young Forests, broken savannas of Senegal, Guinea & Cameroon. Pan troglodytes Ken Lucas Please note that prints are for personal display purposes only

Coke's hartebeests - allogrooming
JW-1312 Coke's hartebeests - allogrooming Alcelaphus buselaphus Ardea London Please note that prints are for personal display purposes only and may not be reproduced in any way

Japanese Macaque JPF 10211 Joshinetsu Kogen National Park, Shiga Highlands, Honshu
JPF-10211 Japanese Macaque Monkeys - group in hot springs to keep warm Joshinetsu Kogen National Park, Shiga Highlands, Honshu

Cheetah - pair grooming each other - Masai Mara National Reserve - Kenya JFL03315
FL-3132 Cheetah - pair grooming each other Maasai Mara National Reserve - Kenya Acinonyx jubatus Ferrero Labat Please note that prints are for personal display purposes only

Cheetah - one female grooming another - Masai Mara National Reserve - Kenya JFL03316
FL-3131 Cheetah - one female grooming another Maasai Mara National Reserve - Kenya Acinonyx jubatus Ferrero Labat Please note that prints are for personal display purposes only

Grey / Common langur - Grooming - Ranthambhore
JVG-3487 Grey / Common langur - Grooming Ranthambhore National Park, India Semnopithecus entellus Alternative spellings: Ranthambhor / Ranthambore / Ranthambor Joanna Van Gruisen Please note that

Japanese Macaque Two grooming a third - Joshinetsu Kogen National Park, Shiga Highlands, Honshu, Japan JPF53071
JPF-14009 Japanese Macaque Two grooming a third Joshinetsu Kogen National Park, Shiga Highlands, Honshu, Japan Macaca fuscata Jean-Paul Ferrero Please note that prints are for personal display

Dingo - Adult biting a juvenile's neck gently, in a lesson in submission, Nadgee Nature Reserve
JPF-13949 Dingo - Adult biting a juvenile's neck gently, in a lesson in submission Nadgee Nature Reserve, far south coast New South Wales

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