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Chimpanzee - adult male showing dominance display - tropical forest - Western Uganda - AfricaSE-60 Chimpanzee - adult male showing dominance display tropical forest - Western Uganda - Africa Pan troglodytes Suzi Eszterhas Please note that prints are for personal display purposes only
Dog. Dog with hair along back raised dominating a puppyJD-21168 Dog. Dog with hair along back raised dominating a puppy John Daniels Please note that prints are for personal display purposes only and may not be reproduced in anyway
Horse WAT 2824 Ponies play fighting. © M. Watson / ARDEA LONDONWAT-2824 Horse - two ponies play fighting M. Watson Please note that prints are for personal display purposes only and may not be reproduced in any way
Wild Grey Wolf - approximately 6 month old pup on left playing with mother - Autumn - Greater Yellowstone EcologicalTOM-1710 Wild Grey Wolf - approximately 6 month old pup on left playing with mother - Autumn Greater Yellowstone Ecological Area - Yellowstone National Park - Wyoming - USA Canis lupus Tom & Pat
Wild Grey Wolf - in snow - approximately 6 month old pup on left showing submission /playfulness to adultsTOM-1691 Wild Grey Wolf - in snow - approximately 6 month old pup on left showing submission /playfulness to adults (creamier wolf is his mother)
Bighorn Sheep Rams - larger ram showing dominance over younger male - Northern Rockies - Alberta - Canada - AutumnTOM-1673 Bighorn Sheep Rams - larger ram showing dominance over younger male Northern Rockies - Alberta - Canada - Autumn Ovis canadensis Tom & Pat Leeson Please note that prints are for personal
DOG. older dog being aggressiveJD-20490 DOG. older dog being aggressive John Daniels Please note that prints are for personal display purposes only and may not be reproduced in any way
Puppies playing (Briard and Jack Russell)JD-20336 Dog - Puppies playing (Briard and Jack Russell) John Daniels Please note that prints are for personal display purposes only and may not be reproduced in any way
DOG Posture. DominantJD-20298 DOG Posture. Dominant John Daniels Please note that prints are for personal display purposes only and may not be reproduced in any way
Polar Bear - x2 fightingWAT-164 POLAR BEAR - two on hind legs, fighting Ursus maritimus M. Watson Please note that prints are for personal display purposes only and may not be reproduced in any way
Eastern / Long-haired Chimpanzee - showing aggression between two chimps - Mahale Mountains National Park - TanzaniaFL-3207 Eastern / Long-haired Chimpanzee - showing aggression between two chimps Mahale Mountains National Park - Tanzania Pan troglodytes schweinfurthi Ferrero Labat Please note that prints are for
Hippopotamus - ritualised fighting, gaping the mouth up to 150 degrees in a show of dominanceFL-3084 Hippopotamus - ritualised fighting, gaping the mouth up to 150 degrees in a show of dominance Maasai Mara National Reserve, Kenya
Southern Elephant Seal FG 268 Valdes Peninsula, Argentina. Mirounga leonina © Fancois Gohier / ARDEA LONDONFG-268 SOUTHERN ELEPHANT SEAL - facing camera Valdes Peninsula, Argentina. Mirounga leonina Francois Gohier Please note that prints are for personal display purposes only
Nile Crocodile - males challenge for dominanceAW-3013 Nile Crocodile - males challenge for dominance Crocodilus niloticus Adrian Warren Please note that prints are for personal display purposes only and may not be reproduced in any way
Dingo - Adult biting a juvenile's neck gently, in a lesson in submission, Nadgee Nature ReserveJPF-13949 Dingo - Adult biting a juvenile's neck gently, in a lesson in submission Nadgee Nature Reserve, far south coast New South Wales
Lion With porcupine. Okavango, Botswana, AfricaCRH-403 LION - WATCHING PORCUPINE Okavango, Botswana, Africa Panthera leo Chris Harvey Please note that prints are for personal display purposes only and may not be reproduced in any way
Wolf / Gray Wolf / Timber Wolf - two males fighting Minnesota USAWAT-14093 Wolf / Gray Wolf / Timber Wolf - two males fighting Minnesota USA Canis lupus M. Watson Please note that prints are for personal display purposes only and may not be reproduced in any way
Japanese Macaque Monkey - two, showing aggression. Hokkaido, JapanWAT-8643 Japanese Macaque Monkey - two, showing aggression Hokkaido, Japan Macaca fuscata M. Watson Please note that prints are for personal display purposes only
Two African Elephants - Bulls spar (mostly play with some dominance overtones)TOM-819 Two African Elephants - Bulls spar (mostly play with some dominance overtones) Along the shore of Lake Kariba, Zimbabwe
Young African Elephant - Bulls engaging in dominance behavior as part of water playTOM-815 Young African Elephant - Bulls engaging in dominance behavior as part of water play. Lake Kariba, Matusadona National Park, Zimbabwe
Stag Beetle 2 males fighting over femalePPG-681 STAG BEETLES - 2 males fighting over female Lucanus cervus Pascal Goetgheluck Please note that prints are for personal display purposes only and may not be reproduced in any way
Borneo Orangutan - old male. Camp Leaky, Tanjung Puting National Park, Borneo, IndonesiaMAR-57 Borneo Orangutan - old male Camp Leaky, Tanjung Puting National Park, Borneo, Indonesia Pongo pygmaeus Thomas Marent Please note that prints are for personal display purposes only
Olive Baboons Young play-fighting Maasai Mara, Kenya, AfricaLA-861 Olive Baboons - Young play fighting Maasai Mara, Kenya, Africa Papio hamadryas anubis Jean Michel Labat Please note that prints are for personal display purposes only
Barbary Apes - Males using newborn to avoid aggression between themJPF-6978 Barbary Macaque / Barbary Ape / Rock Ape - Males using newborn to avoid aggression between them. IUCN Vulnerable Macaca sylvana Calling this species an ape is misleading
"Alpha"-male Proboscis monkey aggressive display deters other males from his harem; after feeding at LabukANZ-1030-m "Alpha"-male Proboscis monkey aggressive display deters other males from his harem after feeding at Labuk Bay Proboscis Monkey Sanctuary
Fallow Deer - young males in combatWAT-979 Fallow DEER - young males in combat Distribution: common throughout Europe. Also found in United Kingdom, Scotland and Northern Ireland. Dama dama M
Black-headed Gull - two immature birds squabblingUSH-857 Black-Headed GULL - Two immature birds squabbling Larus ridibundus Duncan Usher Please note that prints are for personal display purposes only and may not be reproduced in any way
Greenfinch - & Marsh Tit (Parus palustris)USH-798 GREENFINCH - and Marsh Tit (Parus palustris) on ripened sunflower head Carduelis chloris Duncan Usher Please note that prints are for personal display purposes only
Kodiak Bear - 2 animals fighting in lake, Emmen, HollandUSH-3087 Kodiak Bear - 2 animals fighting in lake Ursus arctos middendorffi Duncan Usher Please note that prints are for personal display purposes only and may not be reproduced in any way
European Wolf - 2 animals in snow, one showing submissive behaviour, Bavaria, GermanyUSH-2958 European Wolf - 2 animals in snow, one showing submissive behaviour Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only
European Lynx - play-fightingUSH-262 European Lynx - play fighting captive Felis lynx Duncan Usher Please note that prints are for personal display purposes only and may not be reproduced in any way
Grey Seal - 2 bulls fighting in surf during breeding season. Donna Nook seal sanctuary, Lincolnshire, UKUSH-2552 Grey Seal - 2 bulls fighting in surf during breeding season Donna Nook seal sanctuary, Lincolnshire, UK Halichoerus grypus Duncan Usher Please note that prints are for personal display
European Hare - x2 boxingUSH-202 European HARES - boxing Lepus europaeus Duncan Usher Please note that prints are for personal display purposes only and may not be reproduced in any way
European Wolf - young wolf confronting rank weakest animal, pack dispute Bavaria, GermanyUSH-2006 European Wolf - young wolf confronting rank weakest animal, pack dispute. Note tail position is high of dominant wolf and low on the submissive wolf
European Wolf - Young animal attacking the rank weakest animal in snow, winter Bavaria, GermanyUSH-2002 European Wolf - Young animal attacking the rank weakest animal in snow, winter Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only
European Wolf - pack showing aggresssion towards rank weakest animal Bavaria, GermanyUSH-1975 European Wolf - pack showing aggresssion towards rank weakest animal Bavaria, Germany Canis lupus Duncan Usher Please note that prints are for personal display purposes only
Burchell's / Plains / Common Zebra - stallions, Dominance behavior. Serengeti National Park, Tanzania. May. 3MB778TOM-864 Burchell's / Plains / Common Zebra - stallions, dominance behavior. Serengeti National Park, Tanzania. May. Equus burchelli Tom & Pat Leeson Please note that prints are for personal
Masai Giraffe - Young males "necking" (dominance behavior). East Africa 3mb496TOM-861 Masai Giraffe - Young males "necking" (dominance behavior). Kenya, Africa Giraffa camelopardalis tippelski Tom & Pat Leeson Please note that prints are for personal display purposes
Bighorn Sheep - rams butt heads Northern Rockies. Fall. MS363TOM-1589 Bighorn Sheep - rams butt heads Yellowstone National Park, Northern Rockies. USA, Fall. Ovis canadensis Tom & Pat Leeson Please note that prints are for personal display purposes only
Northern or Steller Sea Lions - bull guarding territory and harem. Pacific Northwest. ML479TOM-1547 Northern / Steller Sealions - bull guarding territory and harem Pacific Northwest, North America Eumetopias jubatus Tom & Pat Leeson Please note that prints are for personal display purposes
sage grouse - two male slap wings (fight) over territory on a lek during spring mating rituals. Western U. S. A B6015TOM-1485 Sage grouse - two male slap wings (fight) over territory on a lek during spring mating rituals. Western U.S.A Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for
Sage Grouse - two male fight over territory on a lek during spring mating rituals. Western U. S. A. BG193TOM-1482 Sage Grouse - two male fight over territory on a lek during spring mating rituals Western U.S.A. Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for personal display
Sage Grouse - two male confront one another over territory on a lek during spring mating season. Western U. S. A. BG313TOM-1481 Sage Grouse - two male confront one another over territory on a lek during spring mating season. Western U.S.A. Centrocercus urophasianus Tom & Pat Leeson Please note that prints are for
Coastal Grizzly Bears wrestling - dominance behavior among big males at salmon fishing areas, McNeil River, AlaskaTOM-1479 Coastal Grizzly Bears wrestling - dominance behavior among big males at salmon fishing areas McNeil River, Alaska
Mountain Goat - two males in a bluff fight, common dominance behavior between goats. Pacific Northwest. MG57TOM-1384 Rocky Mountain Goat - two males in a bluff fight, common dominance behavior between goats Washington, Pacific Northwest
Pronghorn - buck and doe. Buck is chasing doe who has bolted from his harem. Yellowstone National Park, Montana, USATOM-1360 Pronghorn - buck and doe. Buck is chasing doe who has bolted from his harem. Yellowstone National Park, Montana, USA
Bison - bulls sparring Western U. S. A. MB468TOM-1337 Bison - bulls sparring Theodore Roosevelt National Park, North Dakota, U.S.A. Bison bison Tom & Pat Leeson Please note that prints are for personal display purposes only
Bull Bison - fighting, sparring, summer. Theodore Roosevelt National Park, North Dakota. MB249TOM-1335 Bull Bison - fighting, sparring, summer Theodore Roosevelt National Park, North Dakota, USA Bison bison Tom & Pat Leeson Please note that prints are for personal display purposes only